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Plant stem

Plant biology · Stem anatomy & transport

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How does a tree lift water 100 metres into the air without a heart or a single pump?

A plant stem is full of two kinds of tiny pipes. One kind, the xylem, carries water up from the roots, pulled from above as water evaporates from the leaves. The other kind, the phloem, carries sugar made in the leaves to the roots, fruits and growing tips.

A young dicot stem is organised as concentric tissues — epidermis, cortex, a ring of vascular bundles and a central pith — while monocot stems scatter their bundles through ground tissue. Its xylem carries water upward under tension generated by transpiration, and its phloem carries sugar from sources to sinks by osmotically generated pressure flow.

Key facts

Xylem vessel diameter
≈ 20–500 µm (narrow in herbs, widest in ring-porous trees)
Xylem sap velocity
≈ 1–6 m/h in many trees; > 40 m/h in some ring-porous species
Phloem sap velocity
≈ 0.5–1 m/h (typical range 0.3–1.5 m/h)
Phloem sap sucrose
≈ 0.3–0.9 M (10–25% w/v)
Xylem tension in transpiring plants
≈ −0.5 to −3 MPa (lower in drought)
Sieve-element pressure
≈ +0.5 to +2 MPa

Parts you can explore (12)

In the 3D model

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